That is why I stated all areas of calibration wre touched as what you mention IF only the fuel injectors were leaned out a lot then the O2s would see that and just command more fuel.
By keeping the STFTs values within it's operating window then LTFTs are low and PCM does not do anything about it but other areas like timing also have to be tuned and as you notice even in the above testrun and engine taken to 4.700 RPMs the engine only produced 250 HP of the 505 the LS7 outputs.